Research Program & Publications
My research examines palaeoenvironments, palaeoecology, and the life history and evolution of extinct vertebrates. I use a multi-scale, systems approach to disentangle complex interactions between environments and ecosystems in deep time.
(A full list of my publications can be found further down this page)
(A full list of my publications can be found further down this page)
Primary Research
From Organisms to Ecosystems: multi-scale approaches to palaeobiology & environments
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Cretaceous Community Dynamics & Environmental Reconstruction
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Analyzing trends in vertebrate community structure to test the response of species and ecosystems to environmental perturbation. Combination of sedimentary palaeoenvironment data & analyses of fossil assemblages from microsites.
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Applying stable isotope analyses to reconstruct diet, environment, trophic interactions, and habitat preferences in extant and ancient coastal floodplain forest systems.
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Quantifying the variability present in multiple growth proxies found in bone microstructure and reconstructing the growth curves and body size evolution of different dinosaur clades.
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Secondary Research
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Theropod macroevolution & biodiversity, amniote dental histology, the evolution of sexual dimorphism in carnivorans (and particularly pinnipeds)
PDFs are available upon request. Google Scholar page is here
ORCID: 0000-0002-4261-1323
ORCID: 0000-0002-4261-1323
Peer-Reviewed Publications
23. Cullen, TM., Brown, CM., Chiba, K., Brink, KS., Makovicky, PJ., Evans, DC. (Submitted). Growth variability, dimensional scaling, and the interpretation of osteohistological data. Biology Letters.
22. Wilkinson, RD., Cullen, TM., Avrahami, H., Zanno, L., Evans, DC. (Submitted). Paleontology and sedimentology of the Acheroraptor holotype locality, a multi-taxic bonebed in the Hell Creek Formation of Montana, USA. Cretaceous Research.
21. Benson, R., Brown, CM., Campione, N., Cullen, TM., Evans, DC., Zanno, L. (In Review). Response to Schroeder et al: the influence of juvenile dinosaurs on community structure and diversity. Under review at Science.
20. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Huang, L., Evans, DC. (In Review). Dinosaur enamel 13C-enrichment reflects non-analogue environmental conditions in a ‘vanished world’. Under review at Proceedings of the Royal Society B.
19. Cullen, TM., Dunn, R., Suarez, C., Chiba, K. (Accepted). Chp. 2.5: Studying Ecosystems. In The Complete Dinosaur, 3rd Edition. Edited by L. Zanno, T. Holtz, and V. Arbour. Indiana University Press. [invited book chapter contribution]
18. Cullen, TM., Zanno, L., Larson, DW., Todd, E., Currie, PJ., Evans, DC. (In Press). Anatomical, morphometric, and stratigraphic review of theropod biodiversity in the Dinosaur Park Formation (Late Cretaceous; Campanian) of Alberta. Canadian Journal of Earth Sciences (Dale Russell Special Volume).
17. Cullen, TM. (In Press). Stable isotopic analyses of living and extinct crocodylians: implications for understanding their ecology, environments, and physiology. In Crocodylian Biology and Paleobiology. Edited by H Woodward and J Farlow. Indiana University Press. [invited book chapter contribution]
16. Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS., Hu, D., Makovicky, PJ. (2020). Osteohistological analyses reveal diverse strategies of theropod dinosaur body-size evolution. Proceedings of the Royal Society B 287: 20202258. http://dx.doi.org/10.1098/rspb.2020.2258.
15. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Huang, L., Fanti, F., Goodwin, MB., Ryan, MJ., Evans, DC. (2020). Large-scale stable isotope characterization of a Late Cretaceous dinosaur-dominated ecosystem. Geology 48(6): 546-551. (Featured Article & subject of GSA blog)
14. Cullen, TM., Simon, DJ., Benner, EKC., Evans, DC. (2020). Morphology and osteohistology of a large-bodied caenagnathid (Theropoda: Oviraptorosauria) from the Hell Creek Formation (Montana): implications for size-based classifications and growth estimation in theropods. Papers in Palaeontology. https://doi.org/10.1002/spp2/1302
13. Arbour, VM., Evans, DC., Simon, DJ., Cullen, TM., Braman, D. (2019). Cretaceous flora and fauna of the Sustut Group near the Sustut River, northern British Columbia, Canada. Canadian Journal of Earth Sciences. https://doi.org/10.1139/cjes-2019-0031.
12. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Goodwin, MB., Huang, L., and Evans, DC. (2019). Stable isotopic characterization of a coastal floodplain forest community: a case-study for isotopic reconstruction of Mesozoic vertebrate assemblages. Royal Society Open Science 6: 181210. http://dx.doi.org/10.1098/rsos.181210.
11. McFeeters, B., Ryan, MJ., and Cullen, TM. (2018). Response to Brownstein (2018) 'Rebuttal of McFeeters, Ryan and Cullen, 2018'. Vertebrate Anatomy Morphology Palaeontology 6: 73-74. dx.doi.org/10.18435/vamp29343
10. McFeeters, B., Ryan, MJ., and Cullen, TM. (2018). Positional variation in pedal unguals of North American ornithomimids (Dinosauria, Theropoda): a response to Brownstein (2017) . Vertebrate Anatomy Morphology Palaeontology 5. doi: 10.18435/vamp29283
9. LeBlanc, ARH., Brink, K., Cullen, TM., and Reisz, R. (2017). Evolutionary implications of tooth attachment versus tooth implantation: a case study using dinosaur, crocodilian, and mammal teeth. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2017.1354006
8. Evans, DC., Cullen, TM., Larson, DW., and Rego, A. (2017). A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada. Canadian Journal of Earth Sciences. doi: 10.1139/cjes-2017-0034 (Postprint available on PaleorXiv: https://paleorxiv.org/b8mqe/)
7. Cullen, TM. and Evans, DC. (2016). Palaeoenvironmental drivers of vertebrate community composition in the Belly River Group (Campanian) of Alberta, Canada, with implications for dinosaur biogeography. BMC Ecology. doi: 10.1186/s12898-016-0106-8 (Editor's Pick and BMC Ecology top 10 highlight for 2016, and subject of a BMC Series blog)
6. McFeeters, B., Ryan, MJ., Schröder-Adams, C., and Cullen, TM. (2016). A new ornithomimid theropod from the Dinosaur Park Formation of Alberta, Canada. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2016.1221415
5. Cullen, TM., Fanti, F., Capobianco, C., Ryan, MJ., and Evans, DC. (2016). A vertebrate microsite from a marine-terrestrial transition in the Foremost Formation (Campanian) of Alberta, Canada, and the use of faunal assemblage data as a palaeoenvironmental indicator. Palaeogeography, Palaeoclimatology, Palaeoecology 444: 101-114. doi:10.1016/j.palaeo.2015.12.015 (Postprint available on PaleorXiv: https://paleorxiv.org/43j7z/)
4. Cullen, TM., Ryan, MJ., Currie, PJ., Kobayashi, Y., and Evans, DC. (2014). Osteohistological variation in growth marks and osteocyte lacunar density in a theropod dinosaur (Coelurosauria: Ornithomimidae). BMC Evolutionary Biology. doi: 10.1186/s12862-014-0231-y. (Editor’s Pick and a BMC series highlight for November 2014, and included in list of the 12 best figures in BMC Evolutionary Biology for 2014)
3. Evans, DC., Larson, DW., Cullen, TM., & Sullivan, RM. (2014). "Saurornitholestes" robustus is a troodontid (Dinosauria: Theropoda). Canadian Journal of Earth Sciences 51(7): 730-734. doi: 10.1139/cjes-2014-0073
2. Cullen, TM., Fraser, D., Rybczynski, N., and Schroder-Adams, C. (2014). Early evolution of sexual dimorphism and polygyny in Pinnipedia. Evolution 68-5: 1469–1484. (Subject of the May 2014 cover of Evolution) doi: https://dx.doi.org/10.1111/evo.12360 (Postprint available on PaleorXiv: https://paleorxiv.org/zgh62/)
1. Cullen TM., Ryan MJ, Schröder-Adams C, Currie PJ, Kobayashi Y. (2013). An Ornithomimid (Dinosauria) Bonebed from the Late Cretaceous of Alberta, with Implications for the Behavior, Classification, and Stratigraphy of North American Ornithomimids. PLoS ONE 8(3): e58853. doi:10.1371/journal.pone.0058853
22. Wilkinson, RD., Cullen, TM., Avrahami, H., Zanno, L., Evans, DC. (Submitted). Paleontology and sedimentology of the Acheroraptor holotype locality, a multi-taxic bonebed in the Hell Creek Formation of Montana, USA. Cretaceous Research.
21. Benson, R., Brown, CM., Campione, N., Cullen, TM., Evans, DC., Zanno, L. (In Review). Response to Schroeder et al: the influence of juvenile dinosaurs on community structure and diversity. Under review at Science.
20. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Huang, L., Evans, DC. (In Review). Dinosaur enamel 13C-enrichment reflects non-analogue environmental conditions in a ‘vanished world’. Under review at Proceedings of the Royal Society B.
19. Cullen, TM., Dunn, R., Suarez, C., Chiba, K. (Accepted). Chp. 2.5: Studying Ecosystems. In The Complete Dinosaur, 3rd Edition. Edited by L. Zanno, T. Holtz, and V. Arbour. Indiana University Press. [invited book chapter contribution]
18. Cullen, TM., Zanno, L., Larson, DW., Todd, E., Currie, PJ., Evans, DC. (In Press). Anatomical, morphometric, and stratigraphic review of theropod biodiversity in the Dinosaur Park Formation (Late Cretaceous; Campanian) of Alberta. Canadian Journal of Earth Sciences (Dale Russell Special Volume).
17. Cullen, TM. (In Press). Stable isotopic analyses of living and extinct crocodylians: implications for understanding their ecology, environments, and physiology. In Crocodylian Biology and Paleobiology. Edited by H Woodward and J Farlow. Indiana University Press. [invited book chapter contribution]
16. Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS., Hu, D., Makovicky, PJ. (2020). Osteohistological analyses reveal diverse strategies of theropod dinosaur body-size evolution. Proceedings of the Royal Society B 287: 20202258. http://dx.doi.org/10.1098/rspb.2020.2258.
15. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Huang, L., Fanti, F., Goodwin, MB., Ryan, MJ., Evans, DC. (2020). Large-scale stable isotope characterization of a Late Cretaceous dinosaur-dominated ecosystem. Geology 48(6): 546-551. (Featured Article & subject of GSA blog)
14. Cullen, TM., Simon, DJ., Benner, EKC., Evans, DC. (2020). Morphology and osteohistology of a large-bodied caenagnathid (Theropoda: Oviraptorosauria) from the Hell Creek Formation (Montana): implications for size-based classifications and growth estimation in theropods. Papers in Palaeontology. https://doi.org/10.1002/spp2/1302
13. Arbour, VM., Evans, DC., Simon, DJ., Cullen, TM., Braman, D. (2019). Cretaceous flora and fauna of the Sustut Group near the Sustut River, northern British Columbia, Canada. Canadian Journal of Earth Sciences. https://doi.org/10.1139/cjes-2019-0031.
12. Cullen, TM., Longstaffe, FJ., Wortmann, UG., Goodwin, MB., Huang, L., and Evans, DC. (2019). Stable isotopic characterization of a coastal floodplain forest community: a case-study for isotopic reconstruction of Mesozoic vertebrate assemblages. Royal Society Open Science 6: 181210. http://dx.doi.org/10.1098/rsos.181210.
11. McFeeters, B., Ryan, MJ., and Cullen, TM. (2018). Response to Brownstein (2018) 'Rebuttal of McFeeters, Ryan and Cullen, 2018'. Vertebrate Anatomy Morphology Palaeontology 6: 73-74. dx.doi.org/10.18435/vamp29343
10. McFeeters, B., Ryan, MJ., and Cullen, TM. (2018). Positional variation in pedal unguals of North American ornithomimids (Dinosauria, Theropoda): a response to Brownstein (2017) . Vertebrate Anatomy Morphology Palaeontology 5. doi: 10.18435/vamp29283
9. LeBlanc, ARH., Brink, K., Cullen, TM., and Reisz, R. (2017). Evolutionary implications of tooth attachment versus tooth implantation: a case study using dinosaur, crocodilian, and mammal teeth. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2017.1354006
8. Evans, DC., Cullen, TM., Larson, DW., and Rego, A. (2017). A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada. Canadian Journal of Earth Sciences. doi: 10.1139/cjes-2017-0034 (Postprint available on PaleorXiv: https://paleorxiv.org/b8mqe/)
7. Cullen, TM. and Evans, DC. (2016). Palaeoenvironmental drivers of vertebrate community composition in the Belly River Group (Campanian) of Alberta, Canada, with implications for dinosaur biogeography. BMC Ecology. doi: 10.1186/s12898-016-0106-8 (Editor's Pick and BMC Ecology top 10 highlight for 2016, and subject of a BMC Series blog)
6. McFeeters, B., Ryan, MJ., Schröder-Adams, C., and Cullen, TM. (2016). A new ornithomimid theropod from the Dinosaur Park Formation of Alberta, Canada. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2016.1221415
5. Cullen, TM., Fanti, F., Capobianco, C., Ryan, MJ., and Evans, DC. (2016). A vertebrate microsite from a marine-terrestrial transition in the Foremost Formation (Campanian) of Alberta, Canada, and the use of faunal assemblage data as a palaeoenvironmental indicator. Palaeogeography, Palaeoclimatology, Palaeoecology 444: 101-114. doi:10.1016/j.palaeo.2015.12.015 (Postprint available on PaleorXiv: https://paleorxiv.org/43j7z/)
4. Cullen, TM., Ryan, MJ., Currie, PJ., Kobayashi, Y., and Evans, DC. (2014). Osteohistological variation in growth marks and osteocyte lacunar density in a theropod dinosaur (Coelurosauria: Ornithomimidae). BMC Evolutionary Biology. doi: 10.1186/s12862-014-0231-y. (Editor’s Pick and a BMC series highlight for November 2014, and included in list of the 12 best figures in BMC Evolutionary Biology for 2014)
3. Evans, DC., Larson, DW., Cullen, TM., & Sullivan, RM. (2014). "Saurornitholestes" robustus is a troodontid (Dinosauria: Theropoda). Canadian Journal of Earth Sciences 51(7): 730-734. doi: 10.1139/cjes-2014-0073
2. Cullen, TM., Fraser, D., Rybczynski, N., and Schroder-Adams, C. (2014). Early evolution of sexual dimorphism and polygyny in Pinnipedia. Evolution 68-5: 1469–1484. (Subject of the May 2014 cover of Evolution) doi: https://dx.doi.org/10.1111/evo.12360 (Postprint available on PaleorXiv: https://paleorxiv.org/zgh62/)
1. Cullen TM., Ryan MJ, Schröder-Adams C, Currie PJ, Kobayashi Y. (2013). An Ornithomimid (Dinosauria) Bonebed from the Late Cretaceous of Alberta, with Implications for the Behavior, Classification, and Stratigraphy of North American Ornithomimids. PLoS ONE 8(3): e58853. doi:10.1371/journal.pone.0058853
Conference Presentations
(* indicates oral presentation)
38. *Cullen, TM., Larson, DW., Zanno, L., Currie, PJ., Evans, DC. 2020. Theropod biodiversity patterns in the Dinosaur Park Formation (Late Cretaceous: Campanian) of Alberta revealed through morphometrics and biostratigraphy. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
37. *Makovicky, PJ., Darrough, G., Cullen, TM., Fix, M., Gorscak, A., McDonald, A., Saitta, ET., Shinya, A., Stinchcomb, R., Wiersma, J. 2020. New specimens of the basal hadrosauroid, Parrosaurus missouriensis, from the Late Cretaceous of Missouri, USA. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
36. Wilkinson, RD., Cullen, TM., Evans, DC. 2020. Paleontology and sedimentology of a multi-taxic bonebed in the Hell Creek Formation of Montana, USA. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
35. Chinzorig, T., Philips, G., Cullen, TM., Lamb, J., Larson, PL., Rolke, R., Zanno, L. 2020. Ornithomimosaur specimens from the Upper Cretaceous Eutaw Formation (Santonian) of Mississippi: new data on Appalachian theropod dinosaurs. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
34. *Cullen, TM., Larson, DW., Zanno, L., Currie, PJ., Evans, DC. 2020. Morphometrics and biostratigraphy reveal theropod biodiversity patterns in the Dinosaur Park Formation (Late Cretaceous: Campanian) of Alberta. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
33. *Wilkinson, RD., Cullen, TM., Evans, DC. 2020. The diverse faunal assemblage of the Acheroraptor temertyorum holotype locality from the Hell Creek Formation of Montana, USA. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
32. *Arbour, V., Richmond, J., Cullen, TM. 2020. New vertebrate fossil localities in Spatsizi Plateau Wilderness Provincial Park, northern British Columbia. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
31. *Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS, Makovicky, PJ. 2019. Osteohistological variability, growth reconstruction, and trends in the evolution of theropod gigantism. International Symposium on Paleohistology 2019. 31 July - 4 Aug 2019, Cape Town, South Africa.
30. *Cullen, TM. 2019. Specimen Spotlight – the hard life of a young tyrannosaur. Society for the Preservation of Natural History Collections (SPNHC) 2019. 25-31 May 2019, Chicago, IL, USA.
29. *Cullen, TM. 2019. Best practices for destructive sampling in vertebrate paleontology. Society for the Preservation of Natural History Collections (SPNHC) 2019. 25-31 May 2019, Chicago, IL, USA.
28. *Arbour, V., Evans, DC., Simon, DJ., Cullen, TM. 2019. A Maastrichtian-aged leptoceratopsid from the Sustut River, northern BC, and potential for new vertebrate fossil discoveries in the Sustut Basin. Canadian Society of Vertebrate Palaeontology 7th Annual Meeting, 10-13 May 2019. Philip J. Currie Dinosaur Museum, Wembley, Alberta, Canada.
27. *Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS, Makovicky, PJ. 2019. Macroevolutionary patterns of theropod gigantism as revealed through osteohistology and growth curve reconstruction. Canadian Society of Vertebrate Palaeontology 7th Annual Meeting, 10-13 May 2019. Philip J. Currie Dinosaur Museum, Wembley, Alberta, Canada.
26. Arbour, V., Evans, DC., Simon, DJ., Cullen, TM. 2018. Dinosaurs of the Northern Interior Mountains of British Columbia, Canada, including a new high latitude leptoceratopsid. Society of Vertebrate Paleontology 78th Annual Meeting, 17-20 October 2018, Albuquerque, New Mexico, USA.
25. *Cullen, TM. 2018. Testing ecological hypotheses of extant and Cretaceous coastal floodplain forest systems using stable isotope analyses. Society of Vertebrate Paleontology 78th Annual Meeting, 17-20 October 2018, Albuquerque, New Mexico, USA.
24. *Cullen, TM. 2018. Stable isotopic characterization of extant and Cretaceous coastal floodplain forest ecosystems. Canadian Society of Vertebrate Palaeontology 6th Annual Meeting, 14-16 May 2018, Canadian Museum of Nature, Ottawa, Ontario, Canada.
23. *Cullen, TM., Longstaffe, FJ., Wortmann, UG., Goodwin, MB., Huang, L., Evans, DC. 2017. Stable isotope analysis of an extant vertebrate community using palaeontological sampling constraints reveals low ecological resolution in a C3 floodplain system. Society of Vertebrate Paleontology 77th Annual Meeting, 23-26 August 2017, Calgary, Alberta, Canada.
22. *Evans, DC., Cullen, TM., Larson, DW., and Rego, A. 2017. A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada. Canadian Society of Vertebrate Palaeontology 5th Annual Meeting. May 15th-17th, Dinosaur Provincial Park, AB, Canada
21. Cullen, TM., Longstaffe, FJ., Evans, DC. 2016. An intensive multi-taxic stable carbon and oxygen isotopic analysis of vertebrates from a microsite in the Oldman Formation (Late Cretaceous) of Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.26th-29th, Salt Lake City, UT, USA.
20. Benner, EKC., Cullen, TM., Evans, DC. 2016. A new large-bodied caenagnathid specimen (Theropoda, Oviraptorosauria) from the Hell Creek Formation (Late Cretaceous) of Montana, with implications for osteohistological variability in caenagnathids. Society of Vertebrate Paleontology Annual Meeting. Oct.26th-29th, Salt Lake City, UT, USA.
19. *McFeeters, B., Cullen, T. 2016. Holotype of the Late Cretaceous dinosaur Dromiceiomimus brevetertius, with implications for the utility of distal caudal vertebrae in ornithomimid classification. Canadian Paleontological Conference. Aug.26th-28th, Cape Breton University, Sydney, NS, Canada.
18. *Benner, EKC., Cullen, TM., Evans, DC. 2016. Morphological and histological analysis of a new large-bodied caenagnathid specimen (Theropoda: Oviraptorosauria) from the Hell Creek Formation (Montana). Canadian Society of Vertebrate Palaeontology 4th Annual Meeting. May 18th-21st, University of Toronto Mississauga, Mississauga, ON, Canada
17. *Cullen, TM. 2016. The influence of environmental drivers on vertebrate faunal assemblages in the late Cretaceous of Alberta, Canada. Canadian Society of Vertebrate Palaeontology 4th Annual Meeting. May 18th-21st, University of Toronto Mississauga, Mississauga, ON, Canada
16. *Cullen, TM., Evans, DC., Ryan, MJ., and Currie. PJ. 2015. New data on dinosaur faunal turnover and extinction timing in the Dinosaur Park Formation (Late Cretaceous: Campanian) of Alberta, Canada. Society of Vertebrate Paleontology Annual Meeting. Oct.14th-17th, Dallas, TX, USA.
15. Cullen, TM., Evans, DC., Ryan, MJ., Kobayashi, Y., and Currie. PJ. 2014. Variation in intra- and inter-individual osteocyte lacunar density in a theropod dinosaur (Coelurosauria: Ornithomimidae). Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
14. *Leblanc, A., Brink, K., Cullen, TM., Reisz, R. 2014. Tooth sockets and interdental plates: the development and histology of thecodonty in amniotes. Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
13. Larson, D., Cullen, TM., Todd, E., Evans, DC. 2014. Geometric morphometrics of small theropod frontals from the Dinosaur Park formation, Alberta. Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
12. *Cullen, TM. 2014. Intraspecific variation in osteohistologic traits in a size-series of an ornithomimid dinosaur (Theropoda: Coelurosauria). 24th Canadian Paleontology Conference. Aug.28th-31st, Montreal, QC, Canada.
11. Cullen, TM., Ryan, MJ., Capobianco, C., Newbrey, M., and Evans, DC. 2013. A vertebrate microfossil site with a terrestrial-dominated faunal assemblage from the upper Foremost Formation (Campanian) of Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.30th-Nov.2nd, Los Angeles, CA, USA.
10. *Cullen, TM., Fraser, D., Rybczynski, N., and Schröder-Adams, C. 2013. Early evolution of sexual dimorphism and polygyny within Pinnipedia. 23rd Canadian Paleontology Conference. Aug.29th-Sept.1st, Edmonton, AB, Canada.
9. Cullen, TM., Ryan, MJ., Currie, PJ., Kobayashi, Y., and Evans, DC. 2013. Multi- element histological analysis of an ornithomimid (Dinosauria: Theropoda) bonebed from the late cretaceous of Alberta, with implications for skeletochronology in theropods. Second International Symposium on Paleohistology. July 18th-20th, Bozeman, MT, USA.
8. *Cullen, T., Ryan, M., Schröder-Adams, C., Kobayashi, Y., and Currie, P. 2012. Multi- element histological analysis of an ornithomimid (Dinosauria) bone bed from the Horseshoe Canyon Formation, Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.17th-20th, Raleigh, USA.
7. Cullen, TM., Ryan, MJ., Evans, DC., Capobianco, C., and Newbrey, M. 2012. Taxonomy and palaeoecology of a vertebrate microsite from the Foremost Formation of southern Alberta, Canada. 22nd Canadian Paleontology Conference. Sept. 21st-23rd, Toronto, ON, Canada.
6. Cullen, T., Rybczynski, N. and Schröder-Adams, C. 2012. A female Enaliarctos emlongi (Carnivora, Pinnipedimorpha) from the Miocene of western Oregon and the evolution of sexual dimorphism within Pinnipedia. 1st Joint Congress on Evolutionary Biology. July 6th-10th, Ottawa, ON, Canada.
5. *Cullen, T., Ryan, M., Evans, D, Currie, P, and Kobayashi, Y. 2012. Preliminary results of a multi-element histological analysis of an ornithomimid (Dinosauria) bone bed from the late Cretaceous of Alberta. Geological Association of Canada – Mineralogical Association of Canada Meeting. May 27th-29th, St. John’s, NL, Canada.
4. Cullen, T., Pugh, A., Schröder-Adams, C., Halfkenny, B., Rybczynski, N., and Mitchell, T. 2012. Geoheritage Sites of the Canadian Arctic: Increasing Awareness through Education and Co-operation. International Polar Year Conference. April 22nd-27th, Montreal, QC, Canada.
3. Cullen, T., Rybczynski, N., and Schröder-Adams, C. 2011. Description of a small skull specimen of Enaliarctos (Carnivora, Pinnipedimorpha) from the Miocene of western Oregon. Society of Vertebrate Paleontology Annual Meeting. Nov.2nd-5th, Las Vegas, NV, USA.
2. Cullen, T., Rybczynski, N. and Schröder-Adams, C. 2011. Preliminary description of a female Enaliarctos emlongi (Carnivora: Pinnipedia) from the Miocene of western Oregon and implications for the evolution of pinniped mating systems. Canadian Society of Zoologists Annual Meeting. May.16th-20th, Ottawa, ON, Canada.
1. Cullen, T., Ryan, M., Schröder-Adams, C., Kobayashi, Y., Currie, P. 2010. Description of the first ornithomimid (Dinosauria) bonebed from North America with implications for the discrimination, ontogeny and behavior of ornithomimids. Society of Vertebrate Paleontology Annual Meeting. Oct.10th-13th, Pittsburgh, PA, USA.
37. *Makovicky, PJ., Darrough, G., Cullen, TM., Fix, M., Gorscak, A., McDonald, A., Saitta, ET., Shinya, A., Stinchcomb, R., Wiersma, J. 2020. New specimens of the basal hadrosauroid, Parrosaurus missouriensis, from the Late Cretaceous of Missouri, USA. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
36. Wilkinson, RD., Cullen, TM., Evans, DC. 2020. Paleontology and sedimentology of a multi-taxic bonebed in the Hell Creek Formation of Montana, USA. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
35. Chinzorig, T., Philips, G., Cullen, TM., Lamb, J., Larson, PL., Rolke, R., Zanno, L. 2020. Ornithomimosaur specimens from the Upper Cretaceous Eutaw Formation (Santonian) of Mississippi: new data on Appalachian theropod dinosaurs. Society of Vertebrate Palaeontology 80th Annual Meeting, 11-17 October 2020. Virtual Conference.
34. *Cullen, TM., Larson, DW., Zanno, L., Currie, PJ., Evans, DC. 2020. Morphometrics and biostratigraphy reveal theropod biodiversity patterns in the Dinosaur Park Formation (Late Cretaceous: Campanian) of Alberta. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
33. *Wilkinson, RD., Cullen, TM., Evans, DC. 2020. The diverse faunal assemblage of the Acheroraptor temertyorum holotype locality from the Hell Creek Formation of Montana, USA. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
32. *Arbour, V., Richmond, J., Cullen, TM. 2020. New vertebrate fossil localities in Spatsizi Plateau Wilderness Provincial Park, northern British Columbia. Canadian Society of Vertebrate Palaeontology 8th Annual Meeting, 6-7 June 2020. Royal British Columbia Museum, Victoria, BC, Canada. (reviewed and published, though physical conference was cancelled due to covid-19 pandemic)
31. *Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS, Makovicky, PJ. 2019. Osteohistological variability, growth reconstruction, and trends in the evolution of theropod gigantism. International Symposium on Paleohistology 2019. 31 July - 4 Aug 2019, Cape Town, South Africa.
30. *Cullen, TM. 2019. Specimen Spotlight – the hard life of a young tyrannosaur. Society for the Preservation of Natural History Collections (SPNHC) 2019. 25-31 May 2019, Chicago, IL, USA.
29. *Cullen, TM. 2019. Best practices for destructive sampling in vertebrate paleontology. Society for the Preservation of Natural History Collections (SPNHC) 2019. 25-31 May 2019, Chicago, IL, USA.
28. *Arbour, V., Evans, DC., Simon, DJ., Cullen, TM. 2019. A Maastrichtian-aged leptoceratopsid from the Sustut River, northern BC, and potential for new vertebrate fossil discoveries in the Sustut Basin. Canadian Society of Vertebrate Palaeontology 7th Annual Meeting, 10-13 May 2019. Philip J. Currie Dinosaur Museum, Wembley, Alberta, Canada.
27. *Cullen, TM., Canale, JI., Apesteguía, S., Smith, NS, Makovicky, PJ. 2019. Macroevolutionary patterns of theropod gigantism as revealed through osteohistology and growth curve reconstruction. Canadian Society of Vertebrate Palaeontology 7th Annual Meeting, 10-13 May 2019. Philip J. Currie Dinosaur Museum, Wembley, Alberta, Canada.
26. Arbour, V., Evans, DC., Simon, DJ., Cullen, TM. 2018. Dinosaurs of the Northern Interior Mountains of British Columbia, Canada, including a new high latitude leptoceratopsid. Society of Vertebrate Paleontology 78th Annual Meeting, 17-20 October 2018, Albuquerque, New Mexico, USA.
25. *Cullen, TM. 2018. Testing ecological hypotheses of extant and Cretaceous coastal floodplain forest systems using stable isotope analyses. Society of Vertebrate Paleontology 78th Annual Meeting, 17-20 October 2018, Albuquerque, New Mexico, USA.
24. *Cullen, TM. 2018. Stable isotopic characterization of extant and Cretaceous coastal floodplain forest ecosystems. Canadian Society of Vertebrate Palaeontology 6th Annual Meeting, 14-16 May 2018, Canadian Museum of Nature, Ottawa, Ontario, Canada.
23. *Cullen, TM., Longstaffe, FJ., Wortmann, UG., Goodwin, MB., Huang, L., Evans, DC. 2017. Stable isotope analysis of an extant vertebrate community using palaeontological sampling constraints reveals low ecological resolution in a C3 floodplain system. Society of Vertebrate Paleontology 77th Annual Meeting, 23-26 August 2017, Calgary, Alberta, Canada.
22. *Evans, DC., Cullen, TM., Larson, DW., and Rego, A. 2017. A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada. Canadian Society of Vertebrate Palaeontology 5th Annual Meeting. May 15th-17th, Dinosaur Provincial Park, AB, Canada
21. Cullen, TM., Longstaffe, FJ., Evans, DC. 2016. An intensive multi-taxic stable carbon and oxygen isotopic analysis of vertebrates from a microsite in the Oldman Formation (Late Cretaceous) of Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.26th-29th, Salt Lake City, UT, USA.
20. Benner, EKC., Cullen, TM., Evans, DC. 2016. A new large-bodied caenagnathid specimen (Theropoda, Oviraptorosauria) from the Hell Creek Formation (Late Cretaceous) of Montana, with implications for osteohistological variability in caenagnathids. Society of Vertebrate Paleontology Annual Meeting. Oct.26th-29th, Salt Lake City, UT, USA.
19. *McFeeters, B., Cullen, T. 2016. Holotype of the Late Cretaceous dinosaur Dromiceiomimus brevetertius, with implications for the utility of distal caudal vertebrae in ornithomimid classification. Canadian Paleontological Conference. Aug.26th-28th, Cape Breton University, Sydney, NS, Canada.
18. *Benner, EKC., Cullen, TM., Evans, DC. 2016. Morphological and histological analysis of a new large-bodied caenagnathid specimen (Theropoda: Oviraptorosauria) from the Hell Creek Formation (Montana). Canadian Society of Vertebrate Palaeontology 4th Annual Meeting. May 18th-21st, University of Toronto Mississauga, Mississauga, ON, Canada
17. *Cullen, TM. 2016. The influence of environmental drivers on vertebrate faunal assemblages in the late Cretaceous of Alberta, Canada. Canadian Society of Vertebrate Palaeontology 4th Annual Meeting. May 18th-21st, University of Toronto Mississauga, Mississauga, ON, Canada
16. *Cullen, TM., Evans, DC., Ryan, MJ., and Currie. PJ. 2015. New data on dinosaur faunal turnover and extinction timing in the Dinosaur Park Formation (Late Cretaceous: Campanian) of Alberta, Canada. Society of Vertebrate Paleontology Annual Meeting. Oct.14th-17th, Dallas, TX, USA.
15. Cullen, TM., Evans, DC., Ryan, MJ., Kobayashi, Y., and Currie. PJ. 2014. Variation in intra- and inter-individual osteocyte lacunar density in a theropod dinosaur (Coelurosauria: Ornithomimidae). Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
14. *Leblanc, A., Brink, K., Cullen, TM., Reisz, R. 2014. Tooth sockets and interdental plates: the development and histology of thecodonty in amniotes. Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
13. Larson, D., Cullen, TM., Todd, E., Evans, DC. 2014. Geometric morphometrics of small theropod frontals from the Dinosaur Park formation, Alberta. Society of Vertebrate Paleontology Annual Meeting. Nov.5th-8th, Berlin, Germany.
12. *Cullen, TM. 2014. Intraspecific variation in osteohistologic traits in a size-series of an ornithomimid dinosaur (Theropoda: Coelurosauria). 24th Canadian Paleontology Conference. Aug.28th-31st, Montreal, QC, Canada.
11. Cullen, TM., Ryan, MJ., Capobianco, C., Newbrey, M., and Evans, DC. 2013. A vertebrate microfossil site with a terrestrial-dominated faunal assemblage from the upper Foremost Formation (Campanian) of Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.30th-Nov.2nd, Los Angeles, CA, USA.
10. *Cullen, TM., Fraser, D., Rybczynski, N., and Schröder-Adams, C. 2013. Early evolution of sexual dimorphism and polygyny within Pinnipedia. 23rd Canadian Paleontology Conference. Aug.29th-Sept.1st, Edmonton, AB, Canada.
9. Cullen, TM., Ryan, MJ., Currie, PJ., Kobayashi, Y., and Evans, DC. 2013. Multi- element histological analysis of an ornithomimid (Dinosauria: Theropoda) bonebed from the late cretaceous of Alberta, with implications for skeletochronology in theropods. Second International Symposium on Paleohistology. July 18th-20th, Bozeman, MT, USA.
8. *Cullen, T., Ryan, M., Schröder-Adams, C., Kobayashi, Y., and Currie, P. 2012. Multi- element histological analysis of an ornithomimid (Dinosauria) bone bed from the Horseshoe Canyon Formation, Alberta. Society of Vertebrate Paleontology Annual Meeting. Oct.17th-20th, Raleigh, USA.
7. Cullen, TM., Ryan, MJ., Evans, DC., Capobianco, C., and Newbrey, M. 2012. Taxonomy and palaeoecology of a vertebrate microsite from the Foremost Formation of southern Alberta, Canada. 22nd Canadian Paleontology Conference. Sept. 21st-23rd, Toronto, ON, Canada.
6. Cullen, T., Rybczynski, N. and Schröder-Adams, C. 2012. A female Enaliarctos emlongi (Carnivora, Pinnipedimorpha) from the Miocene of western Oregon and the evolution of sexual dimorphism within Pinnipedia. 1st Joint Congress on Evolutionary Biology. July 6th-10th, Ottawa, ON, Canada.
5. *Cullen, T., Ryan, M., Evans, D, Currie, P, and Kobayashi, Y. 2012. Preliminary results of a multi-element histological analysis of an ornithomimid (Dinosauria) bone bed from the late Cretaceous of Alberta. Geological Association of Canada – Mineralogical Association of Canada Meeting. May 27th-29th, St. John’s, NL, Canada.
4. Cullen, T., Pugh, A., Schröder-Adams, C., Halfkenny, B., Rybczynski, N., and Mitchell, T. 2012. Geoheritage Sites of the Canadian Arctic: Increasing Awareness through Education and Co-operation. International Polar Year Conference. April 22nd-27th, Montreal, QC, Canada.
3. Cullen, T., Rybczynski, N., and Schröder-Adams, C. 2011. Description of a small skull specimen of Enaliarctos (Carnivora, Pinnipedimorpha) from the Miocene of western Oregon. Society of Vertebrate Paleontology Annual Meeting. Nov.2nd-5th, Las Vegas, NV, USA.
2. Cullen, T., Rybczynski, N. and Schröder-Adams, C. 2011. Preliminary description of a female Enaliarctos emlongi (Carnivora: Pinnipedia) from the Miocene of western Oregon and implications for the evolution of pinniped mating systems. Canadian Society of Zoologists Annual Meeting. May.16th-20th, Ottawa, ON, Canada.
1. Cullen, T., Ryan, M., Schröder-Adams, C., Kobayashi, Y., Currie, P. 2010. Description of the first ornithomimid (Dinosauria) bonebed from North America with implications for the discrimination, ontogeny and behavior of ornithomimids. Society of Vertebrate Paleontology Annual Meeting. Oct.10th-13th, Pittsburgh, PA, USA.
Non-Refereed Publications
4. Cullen, T. January 10, 2017. Dinosaur distributions in the Belly River. Blog for BMC Series blog. http://blogs.biomedcentral.com/bmcseriesblog/2017/01/10/dinosaur-distributions-belly-river/
3. Cullen, T. July 9, 2016. Microsites and Macroinferences. Blog for 'Valley of the Last Dinosaurs' project of Denver Museum of Nature & Science and the Howard Hughes Medical Institute. http://lastdinos.livesci.org/field-notebook/7/9/microsites-and-macroinferences
2. Cullen, T. July 7, 2016. Field Camps in Many Shapes and Sizes. Blog for 'Valley of the Last Dinosaurs' project of Denver Museum of Nature & Science and the Howard Hughes Medical Institute. http://lastdinos.livesci.org/field-notebook/7/7/field-camps-in-many-shapes-and-sizes
1. Cullen, T. November 19, 2014. Photography in the Field: equal parts business & pleasure. Royal Ontario Museum blogs. https://www.rom.on.ca/en/blog/photography-in-the-field-equal-parts-business-pleasure
3. Cullen, T. July 9, 2016. Microsites and Macroinferences. Blog for 'Valley of the Last Dinosaurs' project of Denver Museum of Nature & Science and the Howard Hughes Medical Institute. http://lastdinos.livesci.org/field-notebook/7/9/microsites-and-macroinferences
2. Cullen, T. July 7, 2016. Field Camps in Many Shapes and Sizes. Blog for 'Valley of the Last Dinosaurs' project of Denver Museum of Nature & Science and the Howard Hughes Medical Institute. http://lastdinos.livesci.org/field-notebook/7/7/field-camps-in-many-shapes-and-sizes
1. Cullen, T. November 19, 2014. Photography in the Field: equal parts business & pleasure. Royal Ontario Museum blogs. https://www.rom.on.ca/en/blog/photography-in-the-field-equal-parts-business-pleasure
Invited Academic Seminars
5. Cullen, TM. ‘Paleobiology Seminar Series’ for the National Museum of Natural History, September 2020, virtual presentation.
4. Cullen, TM. ‘PaleoLunch Seminars’ at the University of Washington / Burke Museum, February 2020, Seattle, WA.
3. Cullen, TM. ‘Collections Forum’ at the Field Museum, December 2018, Chicago, IL.
2. Cullen, TM. 'Atwood Colloquium' at the University of Toronto, April 2015, Toronto, ON.
1. Cullen, TM. 'Advances in Earth Sciences Research Conference, Invited Seminar' at University of Ottawa, March 2014, Ottawa, ON.
4. Cullen, TM. ‘PaleoLunch Seminars’ at the University of Washington / Burke Museum, February 2020, Seattle, WA.
3. Cullen, TM. ‘Collections Forum’ at the Field Museum, December 2018, Chicago, IL.
2. Cullen, TM. 'Atwood Colloquium' at the University of Toronto, April 2015, Toronto, ON.
1. Cullen, TM. 'Advances in Earth Sciences Research Conference, Invited Seminar' at University of Ottawa, March 2014, Ottawa, ON.
Invited Outreach Presentations
9. Cullen, TM. ‘Science Cafe’ for the North Carolina Museum of Natural Sciences, May 2020, virtual presentation.
8. Cullen, TM. ‘Working from home with Mrs. Boltz series’ for the Anne Arundel County Public Schools, March 2020, virtual presentation.
7. Cullen, TM. ‘General Meeting & Holiday Lecture’ at the Earth Science Club of Northern Illinois, December 2019, Glen Ellyn, IL.
6. Cullen, TM. ‘Lunch Hour Talk’ at the Royal British Columbia Museum, August 2019, Victoria, BC.
5. Cullen, TM. ‘21st Annual PaleoFest’ at the Burpee Museum of Natural History, March 2019, Rockford, IL.
4. Cullen, TM. 'A. Watson Armour Seminar Series’ at the Field Museum, December 2018, Chicago, IL.
3. Cullen, TM. ‘Fossil Fest Family Funday’ at the Royal Ontario Museum, January 2017, Toronto, ON.
2. Cullen, TM. ‘Dinosaurs Invade! Weekend’ at the Royal Ontario Museum, January 2015, Toronto, ON.
1. Cullen, TM. 'Students on Ice – Carleton University Antarctic Expedition Lecture Series' at the Canadian Museum of Nature. April 2011, Ottawa, ON
8. Cullen, TM. ‘Working from home with Mrs. Boltz series’ for the Anne Arundel County Public Schools, March 2020, virtual presentation.
7. Cullen, TM. ‘General Meeting & Holiday Lecture’ at the Earth Science Club of Northern Illinois, December 2019, Glen Ellyn, IL.
6. Cullen, TM. ‘Lunch Hour Talk’ at the Royal British Columbia Museum, August 2019, Victoria, BC.
5. Cullen, TM. ‘21st Annual PaleoFest’ at the Burpee Museum of Natural History, March 2019, Rockford, IL.
4. Cullen, TM. 'A. Watson Armour Seminar Series’ at the Field Museum, December 2018, Chicago, IL.
3. Cullen, TM. ‘Fossil Fest Family Funday’ at the Royal Ontario Museum, January 2017, Toronto, ON.
2. Cullen, TM. ‘Dinosaurs Invade! Weekend’ at the Royal Ontario Museum, January 2015, Toronto, ON.
1. Cullen, TM. 'Students on Ice – Carleton University Antarctic Expedition Lecture Series' at the Canadian Museum of Nature. April 2011, Ottawa, ON
Funding, Support, and Collaboration
I would like to thank the following agencies and institutions for their funding and/or support of my research (detailed amounts and further information in my CV PDF):
Natural Sciences and Engineering Research Council of Canada (Alexander Graham Bell Canada Graduate Scholarships)
Government of Ontario (Ontario Graduate Scholarship; Queen Elizabeth II Scholarship for Science and Technology)
Dinosaur Research Institute (project grants, travel funding)
Carleton University (multiple internal scholarships and RA support)
University of Toronto (multiple internal scholarships and RA support)
University of Western Ontario Laboratory for Stable Isotope Science (support, training, and use of analytical equipment)
Canadian Museum of Nature (access to collections, permissions to destructively sample materials)
Royal Ontario Museum (access to collections, use of laboratory facilities, permissions to destructively sample materials)
Royal Tyrrell Museum (access to collections, permissions to destructively sample materials)
Field Museum of Natural History (access to collections, permissions to destructively sample materials)
Kenneth C. Griffin Charitable Fund (support for FMNH postdoctoral position)
Friends of the North Carolina Museum of Natural Sciences (support for NCMNS / NCSU postdoctoral position)
Natural Sciences and Engineering Research Council of Canada (NSERC Postdoctoral Fellowship)
Government of Ontario (Ontario Graduate Scholarship; Queen Elizabeth II Scholarship for Science and Technology)
Dinosaur Research Institute (project grants, travel funding)
Carleton University (multiple internal scholarships and RA support)
University of Toronto (multiple internal scholarships and RA support)
University of Western Ontario Laboratory for Stable Isotope Science (support, training, and use of analytical equipment)
Canadian Museum of Nature (access to collections, permissions to destructively sample materials)
Royal Ontario Museum (access to collections, use of laboratory facilities, permissions to destructively sample materials)
Royal Tyrrell Museum (access to collections, permissions to destructively sample materials)
Field Museum of Natural History (access to collections, permissions to destructively sample materials)
Kenneth C. Griffin Charitable Fund (support for FMNH postdoctoral position)
Friends of the North Carolina Museum of Natural Sciences (support for NCMNS / NCSU postdoctoral position)
Natural Sciences and Engineering Research Council of Canada (NSERC Postdoctoral Fellowship)
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